aqueous
it is wrong of you to imply darwins finches has solved the riddle of evolution, and certainly wrong in implying it has solved the riddle of life
here is a paper from gould that explains why this is so:
http://www.pnas.org/content/94/20/10750.full
Yes I've seen this article before, and read some of the criticisms against Gould that followed. My earlier remark was that the riddle of life was solved by Watson & Crick in their 1953 discovery of DNA. The Darwin's finches did not solve a riddle, they posed one. In the words of the wiki author "these birds were to play an important part in the inception of Darwin's theory of evolution by natural selection". And that's where the riddle was at first solved. Watson & Crick's discovery perfected the theory. (Darwin himself did not classify them as new species--this was the work of another naturalist coincidently named Gould.)
The solution to the riddle of origins is natural genetic modification as culled out under the pressures of natural selection. That's a pretty concise and accurate way to say the riddle has been solved.
It's the other way around - the theory of evolution explains Darwin's finches. The inferred method of propagation is descent with modification. It works whether you assume PE or not. In fact PE is immaterial to this. Genetic modification means more today than it did in Darwin's era, since folks like Mendel and Watson & Crick have expanded its meaning. But Darwin's work is still good in the regard, since he uses the more general term "variation". Without understanding either genetics or molecular biology, Darwin did correctly assess the laws of nature by observing the Darwin's finches. Even that is not in controversy by anything Gould ever wrote. Of that I'm reasonably certain from sampling his works.gould also introduced PE.
both spandrels and PE was introduced because darwins finches and the inferred method of propagation does not explain evolution.
Gould is more interested in mincing the word "gradualism", which is another category of inquiry. His approach was not to dismantle gradualism, but to expand it in such a way that dismantles uniformitarianism. There is some history needed here--uniformitarianism was an earlier rebuttal to catastrophism, which allows God to suspend the laws of nature in order to have creatures appear out of thin air. Uniformitarianism teaches that things progress linearly. It gives the false impression that within a stratum everything is homogeneous. It's now understood that this can't be true, that geological processes which disturb layers prevent uniform fossilization and stratification. Furthermore, fossils are chance occurrences with typically no less than 10,000 years between samples.
Keep in mind that oil exploration has invested heavily in geophysical sciences, that the search for seams which typically produce fossil fuels involves deeper knowledge of what processes led to these formations in the first place, and that this is a field that has grown entirely since Darwin first published.
Before treating PE as a topic in biology, all of the geological non-linearity has to be removed. For math oriented folks, I would compare this to what is known as deconvolution. The idea is that whenever different data sets are merged into one, there are sometimes valid techniques for separating them into their independent original forms. For audiophiles, this something like recovering stereo from mono. In many cases it's next to impossible. This is one reason why you can see so much excruciating detail in articles on paleontology and geology. They can tend to get quite mathematical in their treatments of data for reasons like this.
The riddle of life is one thing. The riddle of how the strata were laid down is quite another. It has been convolved with the riddle of fossilization. The result is that there may appear to be a stretching or compressing of the rate of evolution, when clearly the only thing for certain is that the sample rate is aperiodic. I can't think of how Gould can overlook this, and I assume that he hasn't. It remains to be seen if and when he will give a clear treatment of this, or, if he already has, maybe you or someone else can post a link. In the mean time we have to regard PE as plausible, provided we remove all bias and hyperbole and stay mindful of the caveats. I don't think anyone has ever said that speciation occurs at some fixed rate. It never was a principle of evolution to begin with, so PE changes nothing about the answer to the riddle--unless you are specifically interested in some particular speciation event which still is explained by the same theory.
PE does not contradict the Theory of Evolution, other than to clarify what "gradualism" evidently means. So nothing has really been overturned since the discovery or Darwin's finches. PE might apply to them, but since the theory is was never concerned with the nonlinearity of gradualism, or the anisotropy of the strata, it really doesn't matter too much one way or the other. The fact of genetic modification acted on by natural selection is in no way affected. You might say that the riddle of life is affected, but only if you started out as a uniformitarian, which is probably not the case for most folks who have given it more than a passing thought.
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