Are you responding to me? Or another post?jonmitz said:there are 4 human senses not listed by aristotle (sight, sound, taste, smell, touch)
they are thermoception (heat), nociception (pain), equilbrioception (balance), and proprioception (bodily awareness)
i only know of 3 non human senses: electroception (detect electric fields), magnetoception (detect magnetic fields), and echolocation (similar to sonar)
as to how they evolved, senses did not necessisarily evolve to combat predation. the eye started as a symbiosis of cells that could detect light, and those organisms were able to obtain light energy better and thus did better.
the problem is there is no anwser. also, a problem with your question is that you assume humans are the most advanced species and that evolution was linear directly towards humans. evolution is highly branching, and humans are in no way the only species that are important in this type of question.
another problem with your question is how you would define a sense.
if a microbe is able to use an external fillament to feel around the envrionment, do you consider that equivalent to touch?
if a microbe regects certain chemicals, do you relate that to olfactory or taste (either, doesnt matter)?
it depends on how you define a sense, and how general that definition is or isnt.
in summary, the main problem i think is that you are assuming that humans are the control/omega species.
Are we talking about the beginning of the evolution of senses? Or just in humans?
If we are talking about the origins of sensory perception, then we have to go back to the stimulus-responce perception exhibited in microorganisms. This is microbiology. What are you after? Either way, we have to go back to the ancestorial origins of humans to our amphibian ancestors then to fish then to their sensory perceptions.