Genetics is the answer.
Cold-tolerant plants are capable of activating genes that encode a wide range of enzymes and other proteins that confer cold tolerance. For example:
-- enzymes that are capable of working at low temperatures
-- enzymes that can repair cellular damage caused by frost
-- cytoskeletal-related proteins that can help prevent and repair frost damage
-- proteins that modulate ion channels and signal transduction pathways
-- proteins that prevent damage caused by reactive oxygen species (ROS)
...and more.
http://scholar.google.com.au/scholar?q=cold+tolerance+plants&hl=en&as_sdt=1,5&as_sdtp=on